AboutStaffJohn Lloyd

John Lloyd – Director of Science

2015-08-28-15-37-30John Lloyd joined the Vermont Center for Ecostudies in November 2014. John began his career as an ornithologist at the University of Vermont, where he earned a bachelor’s of science in Wildlife Biology. After a brief stint studying Roseate Terns on Long Island Sound, he headed west. The first stop was the University of Arizona, where John received a master’s of science for his research on the effects of fire suppression on birds in the desert grasslands of southern Arizona. He then spent a year working on a research fellowship in Rocky Mountain National Park, studying the ecological relationships between elk, aspen, and songbirds, before moving to the University of Montana to pursue a doctorate. John’s doctoral research focused on the ecology, evolutionary biology, and conservation of grassland birds in eastern Montana and western North Dakota, with a special focus on Chestnut-collared Longspurs.

After completing his doctorate, John and his family returned to Vermont. His previous research program addressed the ecology and conservation of birds of south Florida and the Caribbean. Since joining VCE, John has focused on creating approaches to research that bridge the spaces between science, policy, and practice and that lead to positive outcomes for nature conservation. He applies these approaches to a variety of topics in a variety of places, from forest fragmentation in Vermont to protected-area management in the Dominican Republic. John also serves as an adjunct associate professor at the University of Vermont.

John’s work as a scientist has been celebrated by the Southwestern Association of Naturalists, who awarded John and his colleagues the 2003 George Miksch Sutton Award in Conservation Research, and by the Florida Ornithological Society, through its Mary J. and William B. Robertson Fellowship Award. In 2017, John was voted an Elective Member of the American Ornithological Society in recognition of his contributions to the study and conservation of birds.

As Director of Science, John strives to ensure that VCE’s science is rigorous and mission-driven; that we are effectively communicating our science with the right people, in the right way, and at the right time; and that VCE’s engagement with citizen-scientists, decision-makers, and other stakeholders is positive, creative, and beneficial to all involved.

Outside of work, John is a runner, fly-fisherman, and devotee of single-speed bikes. He also has a short and mostly undistinguished history as a participant in agricultural fairs.


Visit his ResearchGate page to download or request PDF reprints.

  • Dornelas, M., et al. 2018. BioTIME: a database of biodiversity time series for the Anthropocene. Global Ecology and Biogeography 27:760-786. DOI: 10.1111/geb.12729
  • McFarland, K. P., J. D. Lloyd, S. J. K. Frey, P. L. Johnson, R. B. Chandler, and C. C. Rimmer. 2018. Modeling spatial variation in winter abundance to direct conservation actions for a vulnerable migratory songbird, Bicknell’s Thrush (Catharus bicknelli). Condor 120:517-529.
  • Lloyd, J., S. Garlick, A. Lavallee, and S. Trombulak. 2017. Co-creating an action-oriented agenda for biodiversity research in Vermont. figshare. https://doi.org/10.6084/m9.figshare.5693221.v1
  • Lloyd, J., J. Scarl, J-C Martínez-Sánchez, C. Rimmer, L. Prout, and S. Mathison. 2017. Bicknell’s Thrush mitigation across borders: a strategy for full life-cycle conservation. figshare. https://doi.org/10.6084/m9.figshare.4775680.v1
  • Lloyd, J.D., and C. C. Rimmer. 2017. Surveys of forest birds on Puerto Rico, 2015. Biodiversity Data Journal 5: e20745. https://doi.org/10.3897/BDJ.5.e20745.
  • Rimmer, C.C., P.L. Johnson, and J. D. Lloyd. 2017. Home range size and nocturnal roost locations of Western Chat-Tanagers (Calyptophilus tertius). Wilson Journal of Ornithology 129:611-615.
  • Hill, J. M., and J. D. Lloyd. 2017. A fine-scale U.S. population estimate of a montane spruce-fir bird species of conservation concern. Ecosphere 8(8):e01921. 10.1002/ecs2.1921
  • Lloyd, J.D., C. C. Rimmer, and K. P. McFarland. 2016. Assessing conservation status of resident and migrant birds of Hispaniola with mist-netting. PeerJ 3:31541.DOI: 10.7717/peerj.1541
  • Lloyd, J. D. 2016. Phenotypic variation in Mangrove Cuckoo (Coccyzus minor) across its geographic range. PLoS ONE 11(3): e0152141.doi:10.1371/journal.pone.0152141
  • Lloyd, J. D., and G. L. Slater. 2014. Abundance and distribution of mangrove landbirds in Florida. North American Fauna 80:1-45.
  • Slater, G. L., J. D. Lloyd, and R. W. Snow. 2013. Re-introduction of brown-headed nuthatch and eastern bluebird to South Florida pine rocklands, USA. Pages 63-67 in P. S. Soorae, ed. Global Reintroduction Perspectives 2013. IUCN/SSC, Gland, Switzerland.
  • Slater, G. L., J. D. Lloyd, J. H. Withgott, and K. G. Smith. 2013. Brown-headed Nuthatch (Sitta pusilla), The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. http://bna.birds.cornell.edu/bna/species/349
  • Faccio, S. D., M. Amaral, C. J. Martin, A. Tur, and J. D. Lloyd. 2013. Movement patterns, natal dispersal, and survival of Peregrine Falcons banded in New England. Journal of Raptor Research 47:246-261.
  • Lloyd, J. D. 2013. Mangrove Cuckoo (Coccyzus minor), Neotropical Birds Online (T. S. Schulenberg, Ed.). Ithaca: Cornell Lab of Ornithology. http://neotropical.birds.cornell.edu/portal/species/overview?p_p_spp=202776
  • Lloyd, J. D., G. L. Slater, and J. R. Snyder. 2012. The role of fire-return interval and season of burn on snag dynamics in a south Florida slash-pine forest. Fire Ecology 8:18-31.
  • Lloyd, J. D., and G. L. Slater. 2012. Fire history and the structure of pine-rockland bird assemblages. Natural Areas Journal 32: 247-259.
  • Frieze, R., S. Mullin, and J. D. Lloyd. 2012. Call playbacks increase detection rates of Mangrove Cuckoos. Southeastern Naturalist 11:447-454.
  • Lloyd, J. D. and G. L. Slater. 2011. Influence of fire and water regimes on pineland bird assemblages. Natural Areas Journal 31:270-282.
  • Lloyd, J. D., and T. Doyle. 2011. Abundance and population trends of mangrove landbirds in southwest Florida. Journal of Field Ornithology 82:132-139.
  • Lloyd, J. D., and G. L. Slater. 2011. Abundance and distribution of breeding birds in the pine forests of Grand Bahama, Bahamas. Journal of Caribbean Ornithology 24:1-9.
  • Lloyd, J.D., G. L. Slater, and S. Snow. 2009. Demography of reintroduced Eastern bluebirds and brown-headed nuthatches. Journal of Wildlife Management 73:955-964.
  • Lloyd, J. D., and G. L. Slater. 2007. Environmental factors affecting productivity of brown-headed nuthatches. Journal of Wildlife Management 71:1968-1975.
  • Lloyd, J. D. and J. J. Tewksbury. 2007. Analyzing avian nest survival in forests and grasslands: a comparison of the Mayfield and logistic-exposure methods. Studies in Avian Biology 34:96-104.
  • Tewksbury, J. J., L. Garner, S. Garner, J. D. Lloyd, V. Saab, and T. E. Martin. 2006. Alternative hypotheses of landscape influence on nest predation and brood parasitism in fragmented ecosystems. Ecology 87:759-768.
  • Lloyd, J. D., A. Casey, and M. Trask. 2005. Wildlife hot spots along highways in northwestern Oregon. Pages 680-686 in Proceedings of the 2005 International Conference on Ecology and Transportation.
  • Lloyd, J. D., and T. E. Martin. 2005. Reproductive success of Chestnut-collared Longspurs in native and exotic grassland. Condor 107:365-376.
  • Lloyd, J. D., and T. E. Martin. 2004. Nest-site preference and maternal effects on offspring growth. Behavioral Ecology 15:816-823.
  • Lloyd, J. D. 2004. Partial albinism in a Gray-headed Junco. Colorado Birds 38:114-116.
  • Lloyd, J. D., and T. E. Martin. 2003. Sibling competition and the evolution of developmental rates. Proceedings of the Royal Society of London, Series B 270:735-740. (cover article)
  • Lloyd, J. D., and B. Ruediger. 2003. A rapid assessment process for determining potential wildlife, fish, and plant linkages for highways. Pages 206-226 in Proceedings of the 2003 International Conference on Ecology and Transportation.
  • Kirkpatrick, C., R.W. Mannan, S. DeStefano, and J. D. Lloyd. 2002. Trends in abundance of grassland birds following a spring prescribed burn in southern Arizona. Southwestern Naturalist 47:282-292.
  • Tewksbury, J. J., and J. D. Lloyd. 2001. Positive interactions under nurse plants: spatial scale, stress gradients, and benefactor quality. Oecologia 127:425-434.
  • Lloyd, J.D., R.W. Mannan, S. DeStefano, and C. Kirkpatrick. 1998. The effects of mesquite invasion on a southeastern Arizona grassland bird community. Wilson Bulletin 110:403-408.